College student Chitose Karasuma is determined not to do boring things as she enters the adult world. To this end, this bad-mannered beauty barges into a facility that trains would-be voice actors and actresses, somehow landing a job at “Number One Produce,” a seiyu agency managed by her older brother, Gojo. In Chitose's mind, she's poised for greatness, but finds herself at a loss when she continues to only get minor roles. As she clashes with other girls in the agency, including a cunning airhead and a girl with a Kansai accent, Chitose is about to learn that there's more to succeeding in this competitive industry than she imagined.
High school student Sota is watching the anime Elemental Symphony of Vogelchevalier when he suddenly finds himself drawn into it. When he returns to his world, the heroine of that series, the warrior-princess Selesia Uptiria, comes with him. They soon discover that this is not an isolated case, as several characters from anime, manga, and video games are being transported into what they call "the realm of the gods" by the mysterious girl Military Uniform Princess. Some Creations seek to meet their Creators, while others have more hostile purposes.